Biology & Medicine
Twenty-five centuries of the study of life on one screen — from the humours and the dissection table to evolution, the cell, and the genome.
Each star is a thinker or work; solid lines draw the constellation of a school, dashed threads the passage of ideas between eras.
Select any point on the timeline to read about it.
All entries by era
Ancient Medicine 500 BCE – 500 CE
Hippocrates separates medicine from superstition, Aristotle founds natural history, and Galen builds an anatomy that will rule for a millennium.
- 460 BCE – 370 BCE
The 'father of medicine' sought natural causes of disease rather than divine ones and gave physicians an ethic that survives in the oath.
- 384 BCE – 322 BCE
His careful studies of animals founded natural history and comparative anatomy, and his classification of life stood for two thousand years.
- 129 CE – 216 CE
Physician to gladiators and emperors, his anatomy and physiology dominated Western and Islamic medicine for over a thousand years.
Medieval Medicine 500 CE – 1500 CE
Scholars of the Islamic world preserve and extend Greek medicine, compiling great encyclopaedias and describing the circulation of the blood.
- 980 CE – 1037 CE
His Canon of Medicine organised all known medical knowledge into a system taught in Europe and the Islamic world for six centuries.
- 1025 CE
Avicenna's encyclopaedia codified diagnosis, drugs, and the tenets of medical practice, a standard textbook well into the Renaissance.
- 1213 CE – 1288 CE
He described the pulmonary circulation of the blood three centuries before Europe, correcting Galen from careful reasoning.
Anatomy & the Body 1500 CE – 1670 CE
The Renaissance opens the human body to direct observation: Vesalius corrects Galen and Harvey proves that the heart pumps blood in a circle.
- 1514 CE – 1564 CE
By dissecting human bodies himself, he corrected centuries of Galen's errors and founded modern observational anatomy.
- 1543 CE
Vesalius's magnificently illustrated atlas, published the same year as Copernicus's book, remade anatomy from direct observation.
- 1578 CE – 1657 CE
By measurement and experiment he proved the heart pumps blood around the body in a closed circle, founding modern physiology.
Microscopy & Classification 1670 CE – 1800 CE
The microscope reveals a world of cells and microbes, while Linnaeus gives every living thing a two-word name and a place in a great order.
- 1632 CE – 1723 CE
With hand-ground lenses he first saw bacteria, protozoa, and sperm cells — the founder of microbiology.
- 1635 CE – 1703 CE
Looking at cork through his microscope, he named the 'cell', and his Micrographia revealed a hidden world too small to see.
- 1707 CE – 1778 CE
He gave every species a two-word Latin name and a nested classification, the ordering of life still used by biologists today.
- 1735 CE
Linnaeus's great catalogue placed plants, animals, and minerals into a single hierarchical system of kingdoms, classes, and species.
Cells, Evolution & Germs 1800 CE – 1900 CE
Three great ideas transform biology: all life is made of cells, all species descend by evolution, and disease is caused by microbes.
- 1796 CE
Edward Jenner showed that cowpox protects against deadly smallpox, inventing vaccination and eventually eradicating the disease.
- 1809 CE – 1882 CE
His theory of evolution by natural selection explained the diversity and kinship of all living things from a single simple mechanism.
- 1822 CE – 1884 CE
Breeding pea plants, he discovered the mathematical laws of heredity, founding genetics decades before the world noticed.
- 1822 CE – 1895 CE
He proved that microbes cause fermentation and disease, disproved spontaneous generation, and created vaccines against rabies and anthrax.
- 1839 CE
Schleiden and Schwann proposed that all plants and animals are built of cells — one of biology's great unifying principles.
- 1843 CE – 1910 CE
He identified the microbes of anthrax, tuberculosis, and cholera and set the rigorous rules for proving that a germ causes a disease.
- 1859 CE
Darwin's book laid out the evidence for evolution and natural selection, reshaping biology and our view of humanity's place in nature.
Genetics & Synthesis 1900 CE – 1953 CE
Mendel's laws are rediscovered, chromosomes are shown to carry genes, and evolution and genetics fuse into the modern synthesis.
- 1866 CE – 1945 CE
Working with fruit flies, he showed that genes lie on chromosomes, turning Mendel's abstract factors into physical objects.
- 1928 CE
Alexander Fleming noticed a mould killing bacteria, leading to the first antibiotic and the saving of countless lives.
- 1942 CE
Biologists fused Darwin's natural selection with Mendel's genetics and population mathematics into a single theory of evolution.
The Molecular Age 1953 CE – 2000 CE
The double helix reveals the chemistry of heredity; biologists learn to read, cut, and copy DNA, and set out to sequence a whole genome.
- 1953 CE
Watson and Crick, using Franklin's X-ray images, revealed the structure of DNA and how heredity is copied and passed on.
- 1966 CE
Researchers worked out how triplets of DNA letters spell each amino acid, revealing the universal language shared by all life.
- 1973 CE
The ability to cut and splice genes between organisms launched genetic engineering and the entire biotechnology industry.
Contemporary Biology 2000 CE – 2025 CE
The finished human genome, precise gene editing, and mRNA medicine turn biology into an engineering science of life itself.
- 2003 CE
An international effort read the three billion letters of human DNA, giving medicine and biology a complete genetic reference.
- 2012 CE
A bacterial defence system was turned into a precise tool for editing DNA, making the rewriting of genomes cheap and routine.
- 2020 CE
Decades of molecular biology paid off when mRNA vaccines were designed, tested, and deployed against a new virus within a single year.
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